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Now that we have stated the magnitudes, the properties, and the relative differences of the other internal organs, it remains for us to treat of the organs that contribute to generation. These organs in the female are in all cases internal; in the male they present numerous diversities.

In the blooded animals some males are altogether devoid of testicles, and some have the organ but situated internally; and of those males that have the organ internally situated, some have it close to the loin in the neighbourhood of the kidney and others close to the belly. Other males have the organ situated externally. In the case of these last, the penis is in some cases attached to the belly, whilst in others it is loosely suspended, as is the case also with the testicles; and, in the cases where the penis is attached to the belly, the attachment varies accordingly as the animal is emprosthuretic or opisthuretic.

No fish is furnished with testicles, nor any other creature that has gills, nor any serpent whatever: nor, in short, any animal devoid of feet, save such only as are viviparous within themselves. Birds are furnished with testicles, but these are internally situated, close to the loin. The case is similar with oviparous quadrupeds, such as the lizard, the tortoise and the crocodile; and among the viviparous animals this peculiarity is found in the hedgehog. Others among those creatures that have the organ internally situated have it close to the belly, as is the case with the dolphin amongst animals devoid of feet, and with the elephant among viviparous quadrupeds. In other cases these organs are externally conspicuous.

We have already alluded to the diversities observed in the attachment of these organs to the belly and the adjacent region; in other words, we have stated that in some cases the testicles are tightly fastened back, as in the pig and its allies, and that in others they are freely suspended, as in man.

Fishes, then, are devoid of testicles, as has been stated, and serpents also. They are furnished, however, with two ducts connected with the midriff and running on to either side of the backbone, coalescing into a single duct above the outlet of the residuum, and by 'above' the outlet I mean the region near to the spine. These ducts in the rutting season get filled with the genital fluid, and, if the ducts be squeezed, the sperm oozes out white in colour. As to the differences observed in male fishes of diverse species, the reader should consult my treatise on Anatomy, and the subject will be hereafter more fully discussed when we describe the specific character in each case.

The males of oviparous animals, whether biped or quadruped, are in all cases furnished with testicles close to the loin underneath the midriff. With some animals the organ is whitish, in others somewhat of a sallow hue; in all cases it is entirely enveloped with minute and delicate veins. From each of the two testicles extends a duct, and, as in the case of fishes, the two ducts coalesce into one above the outlet of the residuum. This constitutes the penis, which organ in the case of small ovipara is inconspicuous; but in the case of the larger ovipara, as in the goose and the like, the organ becomes quite visible just after copulation.

The ducts in the case of fishes and in biped and quadruped ovipara are attached to the loin under the stomach and the gut, in betwixt them and the great vein, from which ducts or blood-vessels extend, one to each of the two testicles. And just as with fishes the male sperm is found in the seminal ducts, and the ducts become plainly visible at the rutting season and in some instances become invisible after the season is passed, so also is it with the testicles of birds; before the breeding season the organ is small in some birds and quite invisible in others, but during the season the organ in all cases is greatly enlarged. This phenomenon is remarkably illustrated in the ring-dove and the partridge, so much so that some people are actually of opinion that these birds are devoid of the organ in the winter-time.

Of male animals that have their testicles placed frontwards, some have them inside, close to the belly, as the dolphin; some have them outside, exposed to view, close to the lower extremity of the belly. These animals resemble one another thus far in respect to this organ; but they differ from one another in this fact, that some of them have their testicles situated separately by themselves, while others, which have the organ situated externally, have them enveloped in what is termed the scrotum.

Again, in all viviparous animals furnished with feet the following properties are observed in the testicles themselves. From the aorta there extend vein-like ducts to the head of each of the testicles, and another two from the kidneys; these two from the kidneys are supplied with blood, while the two from the aorta are devoid of it. From the head of the testicle alongside of the testicle itself is a duct, thicker and more sinewy than the other just alluded to-a duct that bends back again at the end of the testicle to its head; and from the head of each of the two testicles the two ducts extend until they coalesce in front at the penis. The duct that bends back again and that which is in contact with the testicle are enveloped in one and the same membrane, so that, until you draw aside the membrane, they present all the appearance of being a single undifferentiated duct. Further, the duct in contact with the testicle has its moist content qualified by blood, but to a comparatively less extent than in the case of the ducts higher up which are connected with the aorta; in the ducts that bend back towards the tube of the penis, the liquid is white-coloured. There also runs a duct from the bladder, opening into the upper part of the canal, around which lies, sheathwise, what is called the 'penis'.

All these descriptive particulars may be regarded by the light of the accompanying diagram; wherein the letter A marks the starting-point of the ducts that extend from the aorta; the letters KK mark the heads of the testicles and the ducts descending thereunto; the ducts extending from these along the testicles are marked MM; the ducts turning back, in which is the white fluid, are marked BB; the penis D; the bladder E; and the testicles XX.

(By the way, when the testicles are cut off or removed, the ducts draw upwards by contraction. Moreover, when male animals are young, their owner sometimes destroys the organ in them by attrition; sometimes they castrate them at a later period. And I may here add, that a bull has been known to serve a cow immediately after castration, and actually to impregnate her.)

So much then for the properties of testicles in male animals.

In female animals furnished with a womb, the womb is not in all cases the same in form or endowed with the same properties, but both in the vivipara and the ovipara great diversities present themselves. In all creatures that have the womb close to the genitals, the womb is two-horned, and one horn lies to the right-hand side and the other to the left; its commencement, however, is single, and so is the orifice, resembling in the case of the most numerous and largest animals a tube composed of much flesh and gristle. Of these parts one is termed the hystera or delphys, whence is derived the word adelphos, and the other part, the tube or orifice, is termed metra. In all biped or quadruped vivipara the womb is in all cases below the midriff, as in man, the dog, the pig, the horse, and the ox; the same is the case also in all horned animals. At the extremity of the so-called ceratia, or horns, the wombs of most animals have a twist or convolution.

In the case of those ovipara that lay eggs externally, the wombs are not in all cases similarly situated. Thus the wombs of birds are close to the midriff, and the wombs of fishes down below, just like the wombs of biped and quadruped vivipara, only that, in the case of the fish, the wombs are delicately formed, membranous, and elongated; so much so that in extremely small fish, each of the two bifurcated parts looks like a single egg, and those fishes whose egg is described as crumbling would appear to have inside them a pair of eggs, whereas in reality each of the two sides consists not of one but of many eggs, and this accounts for their breaking up into so many particles.

The womb of birds has the lower and tubular portion fleshy and firm, and the part close to the midriff membranous and exceedingly thin and fine: so thin and fine that the eggs might seem to be outside the womb altogether. In the larger birds the membrane is more distinctly visible, and, if inflated through the tube, lifts and swells out; in the smaller birds all these parts are more indistinct.

The properties of the womb are similar in oviparous quadrupeds, as the tortoise, the lizard, the frog and the like; for the tube below is single and fleshy, and the cleft portion with the eggs is at the top close to the midriff. With animals devoid of feet that are internally oviparous and viviparous externally, as is the case with the dogfish and the other so-called Selachians (and by this title we designate such creatures destitute of feet and furnished with gills as are viviparous), with these animals the womb is bifurcate, and beginning down below it extends as far as the midriff, as in the case of birds. There is also a narrow part between the two horns running up as far as the midriff, and the eggs are engendered here and above at the origin of the midriff; afterwards they pass into the wider space and turn from eggs into young animals. However, the differences in respect to the wombs of these fishes as compared with others of their own species or with fishes in general, would be more satisfactorily studied in their various forms in specimens under dissection.

The members of the serpent genus also present divergencies either when compared with the above-mentioned creatures or with one another. Serpents as a rule are oviparous, the viper being the only viviparous member of the genus. The viper is, previously to external parturition, oviparous internally; and owing to this perculiarity the properties of the womb in the viper are similar to those of the womb in the selachians. The womb of the serpent is long, in keeping with the body, and starting below from a single duct extends continuously on both sides of the spine, so as to give the impression of thus being a separate duct on each side of the spine, until it reaches the midriff, where the eggs are engendered in a row; and these eggs are laid not one by one, but all strung together. (And all animals that are viviparous both internally and externally have the womb situated above the stomach, and all the ovipara underneath, near to the loin. Animals that are viviparous externally and internally oviparous present an intermediate arrangement; for the underneath portion of the womb, in which the eggs are, is placed near to the loin, but the part about the orifice is above the gut.)

Further, there is the following diversity observable in wombs as compared with one another: namely that the females of horned nonambidental animals are furnished with cotyledons in the womb when they are pregnant, and such is the case, among ambidentals, with the hare, the mouse, and the bat; whereas all other animals that are ambidental, viviparous, and furnished with feet, have the womb quite smooth, and in their case the attachment of the embryo is to the womb itself and not to any cotyledon inside it.

The parts, then, in animals that are not homogeneous with themselves and uniform in their texture, both parts external and parts internal, have the properties above assigned to them.